As plans unfold for a fresh attack on the Neuroi, an enigmatic alien race, the focus turns to capturing Berlin, the pivotal city in the Empire of Karlsland. This strategic move is crucial in eradicating the Neuroi threat across Europe. Yet, with the enemy's ability to swiftly adjust to the battlefield, doubts loom over the effectiveness of the allied forces and the current state of Striker technology. Meanwhile, Yoshika Miyafuji, a Fuso Witch, resumes her medical studies in Lausanne after a recent ordeal left her powerless. Eager to contribute to the war, she awaits the call to arms. Soon, the 501st Joint Fighter Wing must reunite the scattered witches for one last stand against the formidable foe.