Following Cure Muse's successful mission to restore her father's compassionate heart, Ako and Mephisto opt to journey back to Major Land hand in hand. Suddenly, a haunting howl reverberates across the city, stripping away all melodies. Despite their efforts to reach Major Land, Ako and Mephisto are met with silence. Filled with concern, Hibiki and her companions venture to Major Land, only to find a barren realm devoid of its once vibrant music. The populace, resembling emotionless statues, mechanically play instruments bereft of sound. Amidst this eerie silence, Ako's friend Suzu stands as the sole survivor, recounting how Aphrodite, the ruler, inexplicably banished music from their world. Doubtful yet resolute, Ako and Mephisto set out to confront Aphrodite at the palace. Meanwhile, Hibiki, Kanade, and Ellen pursue Suzu, who flees upon Ako's disbelief. Their pursuit leads them to encounter three men intent on seizing the Healing Chest. As revelations unfold, the Pretty Cure must confront the true adversary lurking in the shadows.