In the aftermath of Senki Zesshou Symphogear G, Hibiki Tachibana, Tsubasa Kazanari, and Chris Yukine, guided by Genjuurou Kazanari, resume their duties. While Maria Cadenzavna Eve, Kirika Akatsuki, and Shirabe Tsukuyomi find themselves under protection. Despite a semblance of peace, the emergence of the alchemist Carol Malus Dienheim and her minions, the 'Autoscorers,' threatens to disrupt the tranquility. Carol, wielding a mysterious and formidable power, seeks to unleash chaos and devastation upon the world at any cost. As the six Symphogear wielders confront this enigmatic adversary, they must stand firm to safeguard all they cherish. Yet, the question looms large—will Carol and her Autoscorers prove to be insurmountable obstacles in the battle to preserve the world's destiny?