After the Battle of Britannia and the liberation of Gallia, Yoshika Miyafuji returns to Fuso following her middle school graduation. Her peaceful interlude is shattered when she receives a mysterious letter from her deceased father, unveiling plans for a revolutionary Striker Unit tailored to her magical abilities. As a colossal Neuroi threat emerges in Europe, Yoshika is compelled to join her comrades in the 501st Joint Fighter Wing to confront this new peril. With a dangerous humanoid Neuroi on the horizon and her friends in jeopardy, Yoshika embarks on a perilous journey to the war-torn continent once more.